R. Fraley

British Soldier Lichen

Cladonia cristatella

Brilliant red apothecia resembling tiny soldiers’ caps.
This lichen’s bright scarlet fruiting bodies stand out against its pale green stalks, earning its iconic name “British Soldier.”

British Soldier Lichen on decaying wood British Soldier Lichen on decaying wood
Bright red fruiting bodies atop green podetia.
A classic example of Cladonia cristatella growing on a decaying log in partial sun, often with other crustose lichens.
Size: 4170×2780

Camera: Canon EOS 5D Mk IV
Lens: EF 100mm f/2.8 Macro USM
Settings: ISO 800 • Aperture f/11 • Shutter 1/10 s

Description

Cladonia cristatella, commonly known as the British Soldier Lichen, is a fruticose lichen with erect, stalk-like structures (podetia) that terminate in vivid red apothecia.
The main thallus forms a greenish-gray base layer, from which the upright podetia rise 5–15 mm. These stalks are often branched, with granular or scaly surfaces. The red tips are spore-bearing structures.

This species prefers well-lit, decaying wood, soil, or mossy substrates, particularly in open forests, trailsides, and old stumps. It often coexists with Candelaria, Flavoparmelia, and other pioneer lichens.

Close-up of red apothecia Close-up of red apothecia
Red spore-bearing apothecia atop pale green stalks.
The brilliant pigment derives from anthraquinones, which protect the lichen from UV radiation.
Size: 4170×2780

Camera: Canon EOS 5D Mk IV
Lens: EF 100mm f/2.8 Macro USM
Settings: ISO 400 • Aperture f/8 • Shutter 1/25 s

Behavior

British Soldier lichens grow slowly, often taking years to reach noticeable size. They reproduce via spores from the red apothecia and through fragmentation of the podetia. Typically found in early-successional habitats where competition is minimal.

Color

Podetia: pale green to gray-green from algae and fungal cortex.
Apothecia: intensely scarlet red, forming the distinctive “caps.”

Branching Podetia

Upright branching podetia, typically 5–15 mm tall, rough or granular in texture.

Substrate

Base forms a crustose or squamulose mat, often interwoven with mosses or other lichens.

Size

Entire lichen colony ranges from a few centimeters across; individual stalks 5–15 mm tall and 0.5–1 mm thick. Apothecia typically 0.5–1.5 mm in diameter.

Images

British Soldiers with greenshield lichen British Soldiers with greenshield lichen
Red apothecia rising above a crust of green foliose lichens.
A natural lichen community illustrating how Cladonia species often pioneer new wood surfaces.
Size: 4170×2780

Camera: Canon EOS 5D Mk IV
Lens: EF 100mm f/2.8 Macro USM
Settings: ISO 800 • Aperture f/11 • Shutter 1/8 s